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Support short null for
exon
modepropagate expression char for exons
update feature tests
Examples: expression characters propagated
DRB4*01:03:01N
DRB4*01:03:01:02N/DRB4*01:03:01:13N
DRB4*01:03:01N
DRB4*01:03:01:02N/DRB4*01:03:01:13N
DRB5*01:08:01N/DRB5*01:08:02N
DRB5*01:08:01N/DRB5*01:08:02N
A*01:04:01N
Fixes #161